Type : Venezuela, Bolívar, Gran Sabana, bordering forest near Río Karuai (NW of Mission of santa Teresita de Kavanayén), 1220 m, 26/10/1944, fleurs, Steyermark 59404 (HT:
F0061987F; IT:
NY00312418,
US00136365, VEN27928).
Guyana, Wichabai (Rupununi), savanna, 25-26/10/1937, A.C. Smith 2298 (PT: F).
Brésil, Ceara, 4 km N of Caucaia (Soure), on damp carnauba flat near ocean, 10/06/1945, H. Culter 8366 (PT: F).