Source de la synonymie
Funk, V., Hollowell, T., Berry, P., Kelloff, C. & Alexander, S. N. 2007. Checklist of the Plants of the Guiana Shield (Venezuela: Amazonas, Bolivar, Delta Amacuro; Guyana, Surinam, French Guiana). Contributions from the United States National Herbarium, 55: 1-580. [CD_REF = ]

 

Source de la synonymie
O'Leary, N. & Múlgura, M. E. 2012. A taxonomic revision of the genus Phyla (Verbenaceae). Annals of the Missouri Botanical Garden, 98(4): 578-596. [CD_REF = ]

 

Date de publication : Nov. Gen. Sp. Pl. (quarto ed.) 2: 264-265. 1818 [08/06/1818, fide TL-2, 3143].

Type : "in sylvis Orinocensibus prope Atures et in ripa Apurensi juxta San Fernando".
Venezuela, Amazonas, San Fernando de Atabapo, s.d., Bonpland & Humboldt 1139 (HT: P00604475 [photo: SI]; IT: B-W-11121-01 [photo: SI], P00307284 [photo: SI], P00307283 [photo: SI, F neg.39487]).